Evolution, Old & New Part 21

This has been well extended by Mr. Darwin in a pa.s.sage which begins:--"The affinities of all beings of the same cla.s.s have sometimes been represented by a great tree. I believe that this simile largely speaks the truth."[229]

"What, then," continues Lamarck, "can be the cause of all this? Surely the following: namely, that when individuals of any species change their situation, climate, mode of existence, or habits [conditions of life], their structure, form, organization, and in fact their whole being becomes little by little modified, till in the course of time it responds to the changes experienced by the creature."[230]

In his preface Lamarck had already declared that "the thread which gives us a clue to the causes of the various phenomena of animal organization, in the manifold diversity of its developments, is to be found in the fact that Nature conserves in offspring all that their life and environments has developed in parents." Heredity--"the hidden bond of common descent"--tempered with the modifications induced by changed habits--which changed habits are due to new conditions and surroundings--this with Lamarck, as with Buffon and Dr. Darwin, is the explanation of the diversity of forms which we observe in nature. He now goes on to support this--briefly, in accordance with his design--but with sufficient detail to prevent all possibility of mistake about his meaning.

"In the same climate differences in situation, and a greater or less degree of exposure, affect simply, in the first instance, the individuals exposed to them; but in the course of time, these repeated differences of surroundings in individuals which reproduce themselves continually under similar circ.u.mstances, induce differences which become part of their very nature; so that after many successive generations, these individuals, which were originally, we will say, of any given species, become transformed into a different one."[231]

"Let us suppose that a gra.s.s growing in a low-lying meadow gets carried by some accident to the brow of a neighbouring hill, where the soil is still damp enough for the plant to be able to exist. Let it live here for many generations, till it has become thoroughly accustomed to its position, and let it then gradually find its way to the dry and almost arid soil of a mountain side; if the plant is able to stand the change and to perpetuate itself for many generations, it will have become so changed that botanists will cla.s.s it as a new species."[232]

"The same sort of process goes on in the animal kingdom, but animals are modified more slowly than plants."[233]

The sterility of hybrids, to which Mr. Darwin devotes a great part of the ninth chapter of his 'Origin of Species,'[234] is then touched on--briefly, but sufficiently--as follows:--

"The idea that species were fixed and immutable involved the belief that distinct species could not be fertile _inter se_. But unfortunately observation has proved, and daily proves, that this supposition is unfounded. Hybrids are very common among plants, and quite sufficiently so among animals to show that the boundaries of these so-called immutable species are not so well defined as has been supposed. Often, indeed, there is no offspring between the individuals of what are called distinct species, especially when they are widely different, and again, the offspring when produced is generally sterile; but when there is less difference between the parents, both the difficulty of breeding the hybrid, and its sterility when produced, are found to disappear. In this very power of crossing we see a source from which breeds, and ultimately species, may arise."[235]

Mr. Darwin arrives at the same conclusion. He writes:--

"We must, therefore, either give up the belief of the universal sterility of species when crossed, or we must look at this sterility in animals, not as an indelible characteristic, but as one capable of being removed by domestication.

"Finally, on considering all the ascertained facts on the intercrossing of plants and animals, it may be concluded that some degree of sterility, both in first crosses and in hybrids, is an exceedingly general result, but that it cannot, under our present state of knowledge, be considered as absolutely universal."[236]

Returning to Lamarck, we find him saying:--

"The limits, therefore, of so-called species are not so constant and unvarying as is commonly supposed. Consider also the following. All living forms upon the face of the globe have been brought forth in the course of infinite time by the process of generation only. Nature has directly created none but the lowest organisms; these she is still producing every day, they being, as it were, the first sketches of life, and produced by what is called spontaneous generation. Organs have been gradually developed in these low forms, and these organs have in the course of time increased in diversity and complexity. The power of growth in each living body has given rise to various modes of reproduction, and thus progress, already acquired, has been preserved and handed down to offspring.[237] With sufficient time, favourable conditions of life [_circonstances_], successive changes in the surface of the globe, and the power of new surroundings and habits to modify the organs of living bodies, all animal and vegetable forms have been imperceptibly rendered such as we now see them. It follows that species will be constant only in relation to their environments, and cannot be as old as Nature herself.

"But what are we to say of instinct? Can we suppose that all the tricks, cunning, artifices, precautions, patience, and skill of animals are due to evolution only? Must we not see here the design of an all-powerful Creator? No one certainly will a.s.sign limits to the Creator's power, but it is a bold thing to say that he did not choose to work in this way or that way, when his own handiwork declares to us that this is the way he chose. I find proof in Nature--meaning by nature the _ensemble_ of all that is,[238] but regarding her as herself the effect of an unknown first cause[239]--that she is the author of organization, life, and even sensation; that she has multiplied and diversified the organs and mental powers of the creatures which she sustains and reproduces; that she has developed in animals, through the sole instrumentality of sense of need as establis.h.i.+ng and directing their habits, all actions and all habits, from the simplest up to those which const.i.tute instinct, industry, and finally reason.[240]

"Against this it is alleged that we have no reason to believe species to have changed within any known era. The skeletons of some Egyptian birds, preserved two or three thousand years ago, differ in no particular from the same kind of creatures at the present day. But this is what we should expect, inasmuch as the position and climate of Egypt itself do not appear to have changed. If the conditions of life have not varied, why should the species subjected to those conditions have done so?

Moreover, birds can move about freely, and if one place does not suit them they can find another that does. All that these Egyptian mummies really prove is, that there were animals in Egypt two or three thousand years ago which are like the animals of to-day; but how short a s.p.a.ce is two or three thousand years, as compared with the time which Nature has had at her disposal! A time infinitely great _qua_ man, is still infinitely short _qua_ Nature.[241]

"If, however, we turn to animals under confinement, we find immediate proof that the most startling changes are capable of being produced after some generations of changed habits. In the sixth chapter we shall have occasion to observe the power of changed conditions [_circonstances_] to develop new desires in animals, and to induce new courses of action; we shall see the power which these new actions will have, after a certain amount of repet.i.tion, to engender new habits and tendencies; and we shall also note the effects of use and disuse in either fortifying and developing an organ, or in diminis.h.i.+ng it and causing it to disappear. With plants under domestication, we shall find corresponding phenomena. Species will thus appear to be unchangeable for comparatively short periods only."[242]

It is interesting to see that Mr. Darwin lays no less stress on the study of animals and plants under domestication than Buffon, Dr. Darwin, and Lamarck. Indeed, all four writers appear to have been in great measure led to their conclusions by this very study. "At the commencement of my investigations," writes Mr. Darwin, "it seemed to me probable that a careful study of domesticated animals and of cultivated plants would offer the best chance of making out this obscure problem.

Nor have I been disappointed; in this and in all other perplexing cases, I have invariably found that our knowledge, imperfect though it be, of variation under domestication, afforded the best and safest clue. I may venture to express my conviction of the high value of such studies, though they have been very commonly neglected by naturalists."[243]

In justice to the three writers whom I have named, it should be borne in mind that they also ventured to express their conviction of the high value of these studies. Buffon, indeed, as we have seen, gives animals under domestication the foremost place in his work. He does not treat of wild animals till he has said all he has to say upon our most important domesticated breeds,--on whose descent from one or two wild stocks he is never weary of insisting. It was doubtless because of the opportunities they afforded him for demonstrating the plasticity of living organism that the most important position in his work was a.s.signed to them.

Lamarck professes himself unable to make up his mind about extinct species; how far, that is to say, whole breeds must be considered as having died out, or how far the difference between so many now living and fossil forms is due to the fact that our living species are modified descendants of the fossil ones. Such large parts of the globe were still practically unknown in Lamarck's time, and the recent discovery of the ornithorhynchus has raised such hopes as to what might yet be found in Australia, that he was inclined to think that only such creatures as man found hurtful to him, as, for example, the megatherium and the mastodon, had become truly extinct, nor was he, it would seem, without a hope that these would yet one day be discovered. The climatic and geological changes that have occurred in past ages, would, he believed, account for all the difference which we observe between living and fossil forms, inasmuch as they would have changed the conditions under which animals lived, and therefore their habits and organs would have become correspondingly modified. He therefore rather wondered to find so much, than so little, resemblance between existing and fossil forms.

Buffon took a juster view of this matter; it will be remembered that he concluded his remarks upon the mammoth by saying that many species had doubtless disappeared without leaving any living descendants, while others had left descendants which had become modified.

Lamarck antic.i.p.ated Lyell in supposing geological changes to have been due almost entirely to the continued operation of the causes which we observe daily at work in nature: thus he writes:--

"Every observer knows that the surface of the earth has changed; every valley has been exalted, the crooked has been made straight, and the rough places plain; not even is climate itself stable. Hence changed conditions; and these involve changed needs and habits of life; if such changes can give rise to modifications or developments, it is clear that every living body must vary, especially in its outward character, though the variation can only be perceptible after several generations.

"It is not surprising then that so few living species should be represented in the geologic record. It is surprising rather that we should find any living species represented at all.[244]

"Catastrophes have indeed been supposed, and they are an easy way of getting out of the difficulty; but unfortunately, they are not supported by evidence. Local catastrophes have undoubtedly occurred, as earthquakes and volcanic eruptions, of which the effects can be sufficiently seen; but why suppose any universal catastrophe, when the ordinary progress of nature suffices to account for the phenomena?

Nature is never _brusque_. She proceeds slowly step by step, and this with occasional local catastrophes will remove all our difficulties."[245]

In his fourth chapter Lamarck points out that animals move themselves, or parts of themselves, not through impulsion or movement communicated to them as from one billiard ball to another, but by reason of a cause which excites their irritability, which cause is within some animals and forms part of them, while it is wholly outside of others.[246]

I should again warn the reader to be on his guard against the opinion that any animals can be said to live if they have no "inward motion" of their own which prompts them to act. We cannot call anything alive which moves only as wind and water may make it move, but without any impulse from within to execute the smallest action and without any capacity of feeling. Such a creature does not look sufficiently like the other things which we call alive; it should be first shown to us, so that we may make up our minds whether the facts concerning it have been truly stated, and if so, what it most resembles; we may then cla.s.sify it accordingly.

"Some animals change their place by creeping, some by walking, some by running or leaping; others again fly, while others live in the water and swim.

"The origin of these different kinds of locomotion is to be found in the two great wants of animal life: 1, the means of procuring food; 2, the search after mates with a view to reproduction.

"Since then the power of locomotion was a matter affecting their individual self-preservation, as well as that of their race, the existence of the want led to the means of its being gratified."[247]

Lamarck is practically at one with Dr. Erasmus Darwin, that modification will commonly travel along three main lines which spring from the need of reproduction, of procuring food, and (Dr. Darwin has added) the power of self-protection; but Dr. Darwin's treatment of this part of his subject is more lucid and satisfactory than Lamarck's, inasmuch as he immediately brings forward instances of various modifications which have in each case been due to one of the three main desires above specified, namely, reproduction, subsistence, and self-defence.

Lamarck concludes the chapter with some pa.s.sages which show that he was alive--as what Frenchman could fail to be after Buffon had written?--to the consequences which must follow from the geometrical ratio of increase, and to the struggle for existence, with consequent survival of the fittest, which must always be one of the conditions of any wild animal's existence. The paragraphs, indeed, on this subject are taken with very little alteration from Buffon's work. As Lamarck's theory is based upon the fact that it is on the nature of these conditions that the habits and consequently the structure of any animal will depend, he must have seen that the shape of many of its organs must vary greatly in correlation to the conditions to which it was subjected in the matter of self-protection. I do not see, then, that there is any substantial difference between the positions taken by Dr. Erasmus Darwin and by Lamarck in this respect.

"Let us conclude," he writes, "by showing the means employed by nature to prevent the number of her creatures from injuring the conservation of what has been produced already, and of the general order which should subsist.[248]

"In consequence of the extremely rapid rate of increase of the smaller, and especially of the most imperfect, animals, their numbers would become so great as to prove injurious to the conservation of breeds, and to the progress already made towards more perfect organization, unless nature had taken precautions to keep them down within certain fixed limits which she cannot exceed."[249]

This seems to contain, and in a nutsh.e.l.l, as much of the essence of what Mr. Herbert Spencer and Mr. Charles Darwin have termed the survival of the fittest in the struggle for existence, as was necessary for Lamarck's purpose.

To Lamarck, as to Dr. Darwin and Buffon, it was perfectly clear that the facts, that animals have to find their food under varying circ.u.mstances, and that they must defend themselves in all manner of varying ways against other creatures which would eat them if they could, were simply some of the conditions of their existence. In saying that the surrounding circ.u.mstances--which amount to the conditions of existence--determined the direction in which any plant or animal should be slowly modified, Lamarck includes as a matter of course the fact that the "stronger and better armed should eat the weaker," and thus survive and bear offspring which would inherit the strength and better armour of its parents. Nothing therefore can be more at variance with the truth than to represent Lamarck and the other early evolutionists as ignoring the struggle for existence and the survival of the fittest; these are inevitably implied whenever they use the word "_circonstances_" or environment, as I will more fully show later on, and are also expressly called attention to by the greater number of them.[250]

"Animals, except those which are herbivorous, prey upon one another; and the herbivorous are exposed to the attacks of the flesh-eating races.

"_The strongest and best armed for attack eat the weaker_, and the greater kinds eat the smaller. Individuals of the same race rarely eat one another; they war only with other races than their own."[251]

Dr. Darwin here again has the advantage over Lamarck; for he has pointed out how the males contend with one another for the possession of the females, which I do not find Lamarck to have done, though he would at once have admitted the fact. Lamarck continues:--

"The smaller kinds of animals breed so numerously and so rapidly that they would people the globe to the exclusion of other forms of life, if nature had not limited their inconceivable mult.i.tude. As, however, they are the prey of a number of other creatures, live but a short time, and perish easily with cold, they are kept always within the proportions necessary for the maintenance both of their own and of other races.[252]

"As regards the larger and stronger animals, they would become dominant, and be injurious to the conservation of many other races, if they could multiply in too great numbers. But as it is, they devour one another, and breed but slowly, and few at a birth, so that equilibrium is duly preserved among them. Man alone is the unquestionably dominant animal, but men war among themselves, so that it may be safely said the world will never be peopled to its utmost capacity."[253]

In his fifth chapter Lamarck returns to the then existing arrangement and cla.s.sification of animals.

"Naturalists having remarked that many species, and some genera and even families present characters which as it were isolate them, it has been imagined that these approached or drew further from each other according as their points of agreement or difference seemed greater or less when set down as it were on a chart or map. They regard the small well-marked series which have been styled natural families, as groups which should be placed between the isolated species and their nearest neighbours so as to form a kind of reticulation. This idea, which some of our modern naturalists have held to be admirable, is evidently mistaken, and will be discarded on a profounder and more extended knowledge of organization, and more especially when the distinction has been duly drawn between what is due to the action of special conditions and to general advance of organization."[254]

I take it that Lamarck is here attempting to express what Mr. Charles Darwin has rendered much more clearly in the following excellent pa.s.sage:--

"It should always be borne in mind what sort of intermediate forms must, on the theory [what theory?], have formerly existed. I have found it difficult when looking at any two species to avoid picturing to myself forms _directly_ intermediate between them. But this is a wholly false view; we should always look for forms intermediate between each species and a common but unknown progenitor; and the progenitor will generally have differed in some respects from all its modified descendants. To give a simple ill.u.s.tration: the fantail and pouter pigeons are both descended from the rock pigeon. If we possessed all the intermediate varieties which have ever existed, we should have an extremely close series, between both and the rock pigeon; but we should have no varieties directly intermediate between the fantail and the pouter; none, for instance, combining a tail somewhat expanded with a crop somewhat enlarged, the characteristic features of these two breeds.

These two breeds, moreover, have become so much modified that, if we had no historical or indirect evidence regarding their origin, it would not have been possible to have determined, from a mere comparison of their structure with that of the rock pigeon C. livia, whether they had descended from this species, or from some other allied form, as C.

oenas.

"So with natural species, if we look to forms very distinct--for instance, to the horse and the tapir--we have no reason to suppose that links directly intermediate between them ever existed, but between each and an unknown common parent. The common parent will have had in its whole organization much general resemblance to the tapir and the horse; but in some points of structure it may have differed considerably from both, even perhaps more than they differ from each other. Hence in all such cases we should be unable to recognize the parent form of any two or more species, even if we closely compared the structure of the parent with that of its modified descendants, unless at the same time we had a nearly perfect chain of the intermediate links.

"By the theory of natural selection [surely this is a slip for "by the theory of descent with modification"] all living species have been connected with the parent species of each genus, by differences not greater than we see between the natural and domestic varieties of the same species at the present day; and their parent species, now generally extinct, have in their turn been similarly connected with more ancient forms, and so on backwards, always converging to the common ancestor of each great cla.s.s; so that the number of intermediate and transitional links between all living and extinct species must have been inconceivably great. But a.s.suredly if this theory [the theory of descent with modification or that of "natural selection"?] be true, such have lived upon the earth."[255]