More Letters of Charles Darwin Volume I Part 55

LETTER 360. TO J.D. HOOKER. Down [September 26th 1863].

...About New Zealand, at last I am coming round, and admit it must have been connected with some terra firma, but I will die rather than admit Australia. How I wish mountains of New Caledonia were well worked!...

LETTER 361. TO J.D. HOOKER.

(361/1. In the earlier part of this letter Mr. Darwin refers to a review on Planchon in the "Nat. History Review," April 1865. There can be no doubt, therefore, that "Thomson's article" must be the review of Jordan's "Diagnoses d'especes nouvelles ou meconnues," etc., in the same number, page 226. It deals with "lumpers" and "splitters," and a possible trinomial nomenclature.)

April 17th [1865].

I have been very much struck by Thomson's article; it seems to me quite remarkable for its judgment, force, and clearness. It has interested me greatly. I have sometimes loosely speculated on what nomenclature would come to, and concluded that it would be trinomial. What a name a plant will formally bear with the author's name after genus (as some recommend), and after species and subspecies! It really seems one of the greatest questions which can be discussed for systematic Natural History. How impartially Thomson adjusts the claims of "hair-splitters"

and "lumpers"! I sincerely hope he will pretty often write reviews or essays. It is an old subject of grief to me, formerly in Geology and of late in Zoology and Botany, that the very best men (excepting those who have to write principles and elements, etc.) read so little, and give up nearly their whole time to original work. I have often thought that science would progress more if there was more reading. How few read any long and laborious papers! The only use of publis.h.i.+ng such seems to be as a proof that the author has given time and labour to his work.

LETTER 362. TO J.D. HOOKER. Down, October 22nd and 28th, 1865.

As for the anthropologists being a bete noire to scientific men, I am not surprised, for I have just skimmed through the last "Anthrop.

Journal," and it shows, especially the long attack on the British a.s.sociation, a curious spirit of insolence, conceit, dullness, and vulgarity. I have read with uncommon interest Travers' short paper on the Chatham Islands. (362/1. See Travers, H.H., "Notes on the Chatham Islands," "Linn. Soc. Journ." IX., October 1865. Mr. Travers says he picked up a seed of Edwardsia, evidently washed ash.o.r.e. The stranded logs indicated a current from New Zealand.) I remember your pitching into me with terrible ferocity because I said I thought the seed of Edwardsia might have been floated from Chili to New Zealand: now what do you say, my young man, to the three young trees of the same size on one spot alone of the island, and with the cast-up pod on the sh.o.r.e? If it were not for those unlucky wingless birds I could believe that the group had been colonised by accidental means; but, as it is, it appears by far to me the best evidence of continental extension ever observed. The distance, I see, is 360 miles. I wish I knew whether the sea was deeper than between New Zealand and Australia. I fear you will not admit such a small accident as the wingless birds having been transported on icebergs. Do suggest, if you have a chance, to any one visiting the Islands again, to look out for erratic boulders there. How curious his statement is about the fruit-trees and bees! (362/2. "Since the importation of bees, European fruit-trees and bushes have produced freely." Travers, "Linn. Soc. Journal," IX., page 144.) I wish I knew whether the clover had spread before the bees were introduced...

I saw in the "Gardeners' Chronicle" the sentence about the "Origin"

dying in Germany, but did not know it was by Seemann.

LETTER 363. TO C. LYELL. Down, February 7th [1866].

I am very much obliged for your note and the extract, which have interested me extremely. I cannot disbelieve for a moment Aga.s.siz on Glacial action after all his experience, as you say, and after that capital book with plates which he early published (363/1. "Etudes sur les Glaciers"; Neuchatel, 1840.); as for his inferences and reasoning on the valley of the Amazon that is quite another question, nor can he have seen all the regions to which Mrs. A. alludes. (363/2. A letter from Mrs. Aga.s.siz to Lady Lyell, which had been forwarded to Mr. Darwin. The same letter was sent also to Sir Charles Bunbury, who, in writing to Lyell on February 3rd, 1866, criticises some of the statements. He speaks of Aga.s.siz's observations on glacial phenomena in Brazil as "very astonis.h.i.+ng indeed; so astonis.h.i.+ng that I have very great difficulty in believing them. They shake my faith in the glacial system altogether; or perhaps they ought rather to shake the faith in Aga.s.siz...If Brazil was ever covered with glaciers, I can see no reason why the whole earth should not have been so. Perhaps the whole terrestrial globe was once 'one entire and perfect icicle.'" (From the privately printed "Life" of Sir Charles Bunbury, edited by Lady Bunbury, Volume ii., page 334).) Her letter is not very clear to me, and I do not understand what she means by "to a height of more than three thousand feet." There are no erratic boulders (to which I particularly attended ) in the low country round Rio. It is possible or even probable that this area may have subsided, for I could detect no evidence of elevation, or any Tertiary formations or volcanic action. The Organ Mountains are from six to seven thousand feet in height; and I am only a little surprised at their bearing the marks of glacial action. For some temperate genera of plants, viz., Vaccinium, Andromeda, Gaultheria, Hyperic.u.m, Drosera, Habenaria, inhabit these mountains, and I look at this almost as good evidence of a cold period, as glacial action. That there are not more temperate plants can be accounted for by the isolated position of these mountains. There are no erratic boulders on the Pacific coast north of Chiloe, and but few glaciers in the Cordillera, but it by no means follows, I think, that there may not have been formerly gigantic glaciers on the eastern and more humid side.

In the third edition of "Origin," page 403 (363/3. "Origin," Edition VI., page 335, 1882. "Mr. D. Forbes informs me that he found in various parts of the Cordillera, from lat. 13 deg W. to 30 deg S., at about the height of twelve thousand feet, deeply furrowed rocks...and likewise great ma.s.ses of detritus, including grooved pebbles. Along this whole s.p.a.ce of the Cordillera true glaciers do not now exist, even at much more considerable height. "), you will find a brief allusion, on authority of Mr. D. Forbes, on the former much lower extension of glaciers in the equatorial Cordillera. Pray also look at page 407 at what I say on the nature of tropical vegetation (which I could now much improve) during the Glacial period. (363/4. "During this, the coldest period, the lowlands under the Equator must have been clothed with a mingled tropical and temperate vegetation..." ("Origin," Edition VI., 1882, page 338).)

I feel a strong conviction that soon every one will believe that the whole world was cooler during the Glacial period. Remember Hooker's wonderful case recently discovered of the ident.i.ty of so many temperate plants on the summit of Fernando Po, and on the mountains of Abyssinia.

(363/5. "Dr. Hooker has also lately shown that several of the plants living in the upper parts of the lofty island of Fernando Po, and in the neighbouring Cameroon Mountains, in the Gulf of Guinea, are closely related to those on the mountains of Abyssinia, and likewise to those of temperate Europe" (loc. cit., page 337).) I look at [it] as certain that these plants crossed the whole of Africa from east to west during the same period. I wish I had published a long chapter written in full, and almost ready for the press, on this subject, which I wrote ten years ago. It was impossible in the "Origin" to give a fair abstract.

My health is considerably improved, so that I am able to work nearly two hours a day, and so make some little progress with my everlasting book on domestic varieties. You will have heard of my sister Catherine's easy death last Friday morning. (363/6. Catherine Darwin died in February 1866.) She suffered much, and we all look at her death as a blessing, for there was much fear of prolonged and greater suffering. We are uneasy about Susan, but she has. .h.i.therto borne it better than we could have hoped. (363/7. Susan Darwin died in October 1866.)

Remember glacial action of Lebanon when you speak of no glacial action in S. on Himalaya, and in S.E. Australia.

P.S.--I have been very glad to see Sir C. Bunbury's letter. (363/8. The letter from Bunbury to Lyell, already quoted on this subject. Bunbury writes: "There is nothing in the least NORTHERN, nothing that is not characteristically Brazilian, in the flora of the Organ Mountains.") If the genera which I name from Gardner (363/9. "Travels in the Interior of Brazil," by G. Gardner: London, 1846.) are not considered by him as usually temperate forms, I am, of course, silenced; but Hooker looked over the MS. chapter some ten years ago and did not score out my remarks on them, and he is generally ready enough to pitch into my ignorance and snub me, as I often deserve. My wonder was how any, ever so few, temperate forms reached the mountains of Brazil; and I supposed they travelled by the rather high land and ranges (name forgotten) which stretch from the Cordillera towards Brazil. Cordillera genera of plants have also, somehow, reached the Silla of Caracas. When I think of the vegetation of New Zealand and west coast of South America, where glaciers now descend to or very near to the sea, I feel it rash to conclude that all tropical forms would be destroyed by a considerably cooler period under the Equator.

LETTER 364. TO C. LYELL. Down, Thursday, February 15th [1866].

Many thanks for Hooker's letter; it is a real pleasure to me to read his letters; they are always written with such spirit. I quite agree that Aga.s.siz could never mistake weathered blocks and glacial action; though the mistake has, I know, been made in two or three quarters of the world. I have often fought with Hooker about the physicists putting their veto on the world having been cooler; it seems to me as irrational as if, when geologists first brought forward some evidence of elevation and subsidence, a former Hooker had declared that this could not possibly be admitted until geologists could explain what made the earth rise and fall. It seems that I erred greatly about some of the plants on the Organ Mountains. (364/1. "On the Organ Mountains of Brazil some few temperate European, some Antarctic, and some Andean genera were found by Gardner, which did not exist in the low intervening hot countries"

("Origin," Edition VI., page 336).) But I am very glad to hear about Fuchsia, etc. I cannot make out what Hooker does believe; he seems to admit the former cooler climate, and almost in the same breath to spurn the idea. To retort Hooker's words, "it is inexplicable to me" how he can compare the transport of seeds from the Andes to the Organ Mountains with that from a continent to an island. Not to mention the much greater distance, there are no currents of water from one to the other; and what on earth should make a bird fly that distance without resting many times? I do not at all suppose that nearly all tropical forms were exterminated during the cool period; but in somewhat depopulated areas, into which there could be no migration, probably many closely allied species will have been formed since this period. Hooker's paper in the "Natural History Review" (364/2. Possibly an unsigned article, ent.i.tled "New Colonial Floras" (a review of Grisebach's "Flora of the British West Indian Islands" and Thwaites' "Enumeratio Plantarum Zeylaniae").--"Nat. Hist. Review," January 1865, page 46. See Letter 184.) is well worth studying; but I cannot remember that he gives good grounds for his conviction that certain orders of plants could not withstand a rather cooler climate, even if it came on most gradually.

We have only just learnt under how cool a temperature several tropical orchids can flourish. I clearly saw Hooker's difficulty about the preservation of tropical forms during the cool period, and tried my best to retain one spot after another as a hothouse for their preservation; but it would not hold good, and it was a mere piece of truckling on my part when I suggested that longitudinal belts of the world were cooled one after the other. I shall very much like to see Aga.s.siz's letter, whenever you receive one. I have written a long letter; but a squabble with or about Hooker always does me a world of good, and we have been at it many a long year. I cannot understand whether he attacks me as a wriggler or a hammerer, but I am very sure that a deal of wriggling has to be done.

LETTER 365. TO J.D. HOOKER. Down, July 30th [1866].

Many thanks about the lupin. Your letter has interested me extremely, and reminds me of old times. I suppose, by your writing, you would like to hear my notions. I cannot admit the Atlantis connecting Madeira and Canary Islands without the strongest evidence, and all on that side (365/1. Sir J.D. Hooker lectured on "Insular Floras" at the Nottingham meeting of the British a.s.sociation on August 27th, 1866. His lecture is given in the "Gardeners' Chronicle," 1867, page 6. No doubt he was at this time preparing his remarks on continental extension, which take the form of a judicial statement, giving the arguments and difficulties on both sides. He sums up against continental extension, which, he says, accounts for everything and explains nothing; "whilst the hypothesis of trans-oceanic migration, though it leaves a mult.i.tude of facts unexplained, offers a rational solution of many of the most puzzling phenomena." In his lecture, Sir Joseph wrote that in ascending the mountains in Madeira there is but little replacement of lowland species by those of a higher northern lat.i.tude. "Plants become fewer and fewer as we ascend, and their places are not taken by boreal ones, or by but very few."): the depth is so great; there is nothing geologically in the islands favouring the belief; there are no endemic mammals or batrachians. Did not Bunbury show that some Orders of plants were singularly deficient? But I rely chiefly on the large amount of specific distinction in the insects and land-sh.e.l.ls of P. Santo and Madeira: surely Canary and Madeira could not have been connected, if Madeira and P. Santo had long been distinct. If you admit Atlantis, I think you are bound to admit or explain the difficulties.

With respect to cold temperate plants in Madeira, I, of course, know not enough to form an opinion; but, admitting Atlantis, I can see their rarity is a great difficulty; otherwise, seeing that the lat.i.tude is only a little north of the Persian Gulf, and seeing the long sea-transport for seeds, the rarity of northern plants does not seem to me difficult. The immigration may have been from a southerly direction, and it seems that some few African as well as coldish plants are common to the mountains to the south.

Believing in occasional transport, I cannot feel so much surprise at there being a good deal in common to Madeira and Canary, these being the nearest points of land to each other. It is quite new and very interesting to me what you say about the endemic plants being in so large a proportion rare species. From the greater size of the workshop (i.e., greater compet.i.tion and greater number of individuals, etc.) I should expect that continental forms, as they are occasionally introduced, would always tend to beat the insular forms; and, as in every area, there will always be many forms more or less rare tending towards extinction, I should certainly have expected that in islands a large proportion of the rarer forms would have been insular in their origin. The longer the time any form has existed in an island into which continental forms are occasionally introduced, by so much the chances will be in favour of its being peculiar or abnormal in nature, and at the same time scanty in numbers. The duration of its existence will also have formerly given it the best chance, when it was not so rare, of being widely distributed to adjoining archipelagoes. Here is a wriggle: the older a form is, the better the chance will be of its having become developed into a tree! An island from being surrounded by the sea will prevent free immigration and compet.i.tion, hence a greater number of ancient forms will survive on an island than on the nearest continent whence the island was stocked; and I have always looked at Clethra (365/2. Clethra is an American shrubby genus of Ericaceae, found nowhere nearer to Madeira than North America. Of this plant and of Persea, Sir Charles Lyell ("Principles," 1872, Volume II., page 422) says: "Regarded as relics of a Miocene flora, they are just such forms as we should naturally expect to have come from the adjoining Miocene continent." See also "Origin of Species," Edition VI., page 83, where a similar view is quoted from Heer.) and the other extra-European forms as remnants of the Tertiary flora which formerly inhabited Europe. This preservation of ancient forms in islands appears to me like the preservation of ganoid fishes in our present freshwaters. You speak of no northern plants on mountains south of the Pyrenees: does my memory quite deceive me that Boissier published a long list from the mountains in Southern Spain? I have not seen Wollaston's, "Catalogue," (365/4. Probably the "Catalogue of the Coleopterous Insects of the Canaries in the British Museum,"

1864.) but must buy it, if it gives the facts about rare plants which you mention.

And now I have given more than enough of my notions, which I well know will be in flat contradiction with all yours.

Wollaston, in his "Insecta Maderensia" (365/5. "Insecta Maderensia,"

London, 1854.), 4to, page 12, and in his "Variation of Species," pages 82-7, gives the case of apterous insects, but I remember I worked out some additional details.

I think he gives in these same works the proportion of European insects.

LETTER 366. TO J.D. HOOKER.

(366/1. Sir Joseph had asked (July 31st, 1866): "Is there an evidence that the south of England and of Ireland were not submerged during the Glacial epoch, when the W. and N. of England were islands in a glacial sea? And supposing they were above water, could the present Atlantic and N.W. of France floras we now find there have been there during the Glacial epoch?--Yet this is what Forbes demands, page 346. At page 347 he sees this objection, and wriggles out of his difficulty by putting the date of the Channel 'towards the close of the Glacial epoch.' What does Austen make the date of the Channel?--ante or post Glacial?" The changes in level and other questions are dealt with in a paper by R.A.C.

Austen (afterwards G.o.dwin-Austen), "On the Superficial Acc.u.mulations of the Coasts of the English Channel and the Changes they indicate."

"Quart. Journ. Geol. Soc." VII., 1851, page 118. Obit. notice by Prof.

Bonney in the "Proc. Geol. Soc." XLI., page 37, 1885.)

Down, August 3rd [1866].

I will take your letter seriatim. There is good evidence that S.E.

England was dry land during the Glacial period. I forget what Austen says, but Mammals prove, I think, that England has been united to the Continent since the Glacial period. I don't see your difficulty about what I say on the breaking of an isthmus: if Panama was broken through would not the fauna of the Pacific flow into the W. Indies, or vice versa, and destroy a mult.i.tude of creatures? Of course I'm no judge, but I thought De Candolle had made out his case about small areas of trees.

You will find at page 112, 3rd edition "Origin," a too concise allusion to the Madeira flora being a remnant of the Tertiary European flora.

I shall feel deeply interested by reading your botanical difficulties against occasional immigration. The facts you give about certain plants, such as the heaths, are certainly very curious. (366/2. In Hooker's lecture he gives St. Dabeoc's Heath and Calluna vulgaris as the most striking of the few boreal plants in the Azores. Darwin seems to have been impressed by the boreal character of the Azores, thus taking the opposite view to that of Sir Joseph. See Letter 370, note.) I thought the Azores flora was more boreal, but what can you mean by saying that the Azores are nearer to Britain and Newfoundland than to Madeira?--on the globe they are nearly twice as far off. (366/3. See Letter 368.) With respect to sea currents, I formerly made enquiries at Madeira, but cannot now give you the results; but I remember that the facts were different from what is generally stated: I think that a s.h.i.+p wrecked on the Canary Islands was thrown up on the coast of Madeira.

You speak as if only land-sh.e.l.ls differed in Madeira and Porto Santo: does my memory deceive me that there is a host of representative insects?

When you exorcise at Nottingham occasional means of transport, be honest, and admit how little is known on the subject. Remember how recently you and others thought that salt water would soon kill seeds.

Reflect that there is not a coral islet in the ocean which is not pretty well clothed with plants, and the fewness of the species can hardly with justice be attributed to the arrival of few seeds, for coral islets close to other land support only the same limited vegetation. Remember that no one knew that seeds would remain for many hours in the crops of birds and retain their vitality; that fish eat seeds, and that when the fish are devoured by birds the seeds can germinate, etc. Remember that every year many birds are blown to Madeira and to the Bermudas. Remember that dust is blown 1,000 miles over the Atlantic. Now, bearing all this in mind, would it not be a prodigy if an unstocked island did not in the course of ages receive colonists from coasts whence the currents flow, trees are drifted and birds are driven by gales. The objections to islands being thus stocked are, as far as I understand, that certain species and genera have been more freely introduced, and others less freely than might have been expected. But then the sea kills some sorts of seeds, others are killed by the digestion of birds, and some would be more liable than others to adhere to birds' feet. But we know so very little on these points that it seems to me that we cannot at all tell what forms would probably be introduced and what would not. I do not for a moment pretend that these means of introduction can be proved to have acted; but they seem to me sufficient, with no valid or heavy objections, whilst there are, as it seems to me, the heaviest objections on geological and on geographical distribution grounds (pages 387, 388, "Origin" (366/4. Edition III., or Edition VI., page 323.) to Forbes'

enormous continental extensions. But I fear that I shall and have bored you.

LETTER 367. J.D. HOOKER TO CHARLES DARWIN.

(367/1. In a letter of July 31st, Sir J.D. Hooker wrote, "You must not suppose me to be a champion of continental connection, because I am not agreeable to trans-oceanic migration...either hypothesis appears to me well to cover the facts of oceanic floras, but there are grave objections to both, botanical to yours, geological to Forbes'.")

The following interesting letters give some of Sir Joseph's difficulties.)

Kew, August 4th, 1866.

You mention ("Journal") no land-birds, except introduced, upon St.

Helena. Beatson (Introduction xvii) mentions one (367/2. Aegialitis sanctae-helenae, a small plover "very closely allied to a species found in South Africa, but presenting certain differences which ent.i.tle it to the rank of a peculiar species" (Wallace, "Island Life," page 294).

In the earlier editions of the "Origin" (e.g. Edition III., page 422) Darwin wrote that "Madeira does not possess one peculiar bird."

In Edition IV., 1866, page 465, the mistake was put right.) "in considerable numbers," resembles sand-lark--is called "wire bird," has long greenish legs like wires, runs fast, eyes large, bill moderately long, is rather shy, does not possess much powers of flight. What was it? I have written to ask Sclater, also about birds of Madeira and Azores. It is a very curious thing that the Azores do not contain the (non-European) American genus Clethra, that is found in Madeira and Canaries, and that the Azores contain no trace of American element (beyond what is common to Madeira), except a species of Sanicula, a genus with hooked bristles to the small seed-vessels. The European Sanicula roams from Norway to Madeira, Canaries, Cape Verde, Cameroons, Cape of Good Hope, and from Britain to j.a.pan, and also is, I think, in N. America; but does not occur in the Azores, where it is replaced by one that is of a decidedly American type.

This tells heavily against the doctrine that joins Atlantis to America, and is much against your trans-oceanic migration--for considering how near the Azores are to America, and in the influence of the Gulf-stream and prevalent winds, it certainly appears marvellous. Not only are the Azores in a current that sweeps the coast of U. States, but they are in the S.W. winds, and in the eye of the S.W. hurricanes!