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There is a cerebral form peculiar to man and the apes; and so in the cerebral convolutions, wherever they appear, there is a general unity of arrangement, a plan, the type of which is common to all these creatures." Professor Huxley says: "It is most remarkable that, as soon as all the princ.i.p.al sulci appear, the pattern according to which they are arranged is identical with the corresponding sulci in man. The surface of the brain of the monkey exhibits a sort of skeleton map of man's, and in the man-like apes the details become more and more filled in, until it is only in minor characters that the chimpanzee's or orang's brain can be structurally distinguished from man's."
The facts of anatomy, at least, are all against us. Struggle as we may, be as sn.o.bbish as we will, we cannot shake off these poor relations of ours. Our adult anatomy at once betrays our ancestry, if we attempt to deny it. Read the first chapter of that remarkable book by Professor Drummond on the "Ascent of Man," the chapter on the ascent of the body, and the second chapter on the scaffolding left in the body. The tips of our ears and our rudimentary ear muscles, the hair on hand and arm, and the little plica semilunaris, or rudimentary third eyelid in the inner angle of our eyes, the vermiform appendage of the intestine, the coracoid process on our shoulder-blades, the atlas vertebra of our necks--to say nothing of the coccyx at the other end of the backbone--many malformations, and a host of minor characteristics all refute our denial.
If we appeal from adult anatomy to embryology the case becomes all the worse for us. Our ear is lodged in the gill-slit of a fish, our jaws are branchial arches, our hyoid bone the rudiment of this system of bones supporting the gills. Our circulation begins as a veritable fish circulation; our earliest skeleton is a notochord; Meckel's cartilage, from which our lower jaw and the bones of our middle ear develop, is a whole genealogical tree of disagreeable ancestors. Our glandula thyreoidea has, according to good authorities, an origin so slimy that it should never be mentioned in polite society. The origin of our kidneys appears decidedly vermian.
Time fails me to read merely the name of the witnesses which could be summoned from our own bodies to witness against us.
Even if the testimony of some of these witnesses is not as strong as many think, and we have misunderstood several of them, they are too numerous and their stories hang too well together not to impress an intelligent and impartial jury. But what if it is all true? What if, as some think, our millionth cousin, the tiger or cat, is anatomically a better mammal than I? His teeth and claws and magnificent muscles are of small value compared with man's mental power.
What a comedy that man should work so hard to prove that his chief glory is his opposable thumb, or a few ounces of brain matter! Man's glory is his mind and will, his reason and moral powers, his vision of, and communion with, G.o.d. And supposing it be true, as I believe it is true, that the animal has the germ of these also, does that cloud my mind or obscure my vision or weaken my action? It bids me only strive the harder to be worthy of the n.o.ble ancestors who have raised me to my higher level and on whose buried shoulders I stand.
Whatever may have been our origin, whoever our ancestors, we are men. Then let us play the man. If we will but play our part as well as our old ancestors played theirs, if we will but walk and act according to our light one-half as heroically and well as they groped in the darkness, we need not worry about the future. That will be a.s.sured.
Says Professor Huxley: "Man now stands as on a mountain-top far above the level of his humble fellows, and transfigured from his grosser nature by reflecting here and there a ray from the infinite source of truth. And thoughtful man, once escaped from the blinding influences of traditional prejudice, will find in the lowly stock whence man has sprung the best evidence of the splendor of his capacities, and will discern in his long progress through the past a reasonable ground of faith in his attainment of a n.o.bler future."
We have sketched hastily and in rude outline the anatomical structure of the successive stages of man's ancestry; let us now, in a very brief recapitulation, condense this chronicle into a historical record of progress.
We began with the amoeba. This could not have been the beginning.
In all its structure it tells us of something earlier and far simpler, but what this earlier ancestor was we do not know. Rather more highly organized relatives of the amoeba, the flagellata, have produced a membrane, and swim by means of vibratile, whiplash-like flagella. We must emphasize that these little animals correspond in all essential respects to the cells of our bodies; they are unicellular animals. And the cell once developed remains essentially the same structure, modified only in details, throughout higher animals. And these unicellular animals have the rudiments of all our functions. Their protoplasm and functions seem to differ from those of higher animals only in degree, not in kind. And the more we consider both these facts the more remarkable and suggestive do they become.
Cells with membranes can unite in colonies capable of division of labor and differentiation. And magosphaera is just such a little spheroidal colony. But the cells are still all alike, each one performs all functions equally well. But in volvox division of labor and differentiation of structure have taken place. Certain cells have become purely reproductive, while the rest gather nutriment for these, but are at the same time sensitive and locomotive, excretory and respiratory. The first function to have cells specially devoted to it is the reproductive; this is a function absolutely necessary for the maintenance of the species. For the nutritive cells die when they have brought the reproductive cells to their full development.
These few nutritive cells represent the body of all higher animals in contrast with the reproductive elements. And with the development of a body, death, as a normal process, enters the world. The dominant function is here evidently the reproductive, and the whole body is subservient to this.
In hydra the union and differentiation of cells is carried further.
But the cells are still much alike and only slowly lose their own individuality in that of the whole animal. This is shown in the fact that each entodermal cell digests its own particles of food, although the nutriment once digested diffuses to all parts of the body. Also almost any part of the animal containing both ectoderm and entoderm can be cut off and will develop into a new animal.
But beside the reproductive cells and tissues hydra has developed a very simple digestive system, in which the newly caught food at least macerates and begins to be dissolved. This is the second essential function. The animal can, and the plant as a rule does, exist with only the lowest rudiments of anything like nervous or muscular power; but no species can exist without good powers of digestion and reproduction. These essential organs must first develop and the higher must wait. And the inner, digestive, layer of cells persists in our bodies as the lining of the mid-intestine. We compared hydra therefore to a little patch of the lining of our intestine covered with a flake of epidermis; only these layers in hydra possess powers lost to the corresponding cells of our bodies in the process of differentiation. Notice, please, that when cell or organ has once been developed it persists, as a rule, modified, but not lost. Nature's experiments are not in vain; her progress is very slow but sure. But hydra has also the promise of better things, traces of muscular and nervous tissue. There are still no compact muscles, like our own, much less ganglion or brain or nerve-centre of individuality. The tissues are diffuse, but they are the materials out of which the organs of higher animals will crystallize, so to speak. Notice also that these higher muscles and nerves are here entirely subservient to, and exist for, digestion and reproduction.
In the turbellaria the reproductive system has reached a very high grade of development. It is a complex and beautifully constructed organ. The digestive system has also vastly improved; it has its own muscular layers, and often some means of grasping food. But it is slower in reaching its full development than the reproductive system. But all the muscles are no longer attached to the stomach; they are beginning to a.s.sert their independence, and, in a rude way, to build a body-wall. But they are in many layers, and run in almost all directions. Some of these layers will disappear, but the most important ones, consisting of longitudinal and transverse fibres, will persist in higher forms. Locomotion by means of these muscles is slowly coming into prominence. They are no longer merely slaves of digestion.
But a muscular fibril contracts only under the stimulus of a nervous impulse. More nerve-cells are necessary to control these more numerous muscular fibrils. The animal now moves with one end foremost, and that end first comes in contact with food, hindrances, or injurious surroundings. Here the sensory cells of feeling and their nerve fibrils multiply. Remember that these neuro-epithelial sensory cells are suited to respond not merely to pressure, but to a variety of the stimuli, chemical, molecular, and of vibration, which excite our organs of smell, taste, and hearing. Such organs and the directive eyes appear mainly at this anterior end. But a ganglion cell sends an impulse to a muscle because it has received one along a sensory nerve from one or more of these sensory cells. Hence the ganglion cells will increase in number. The old cobweb-like plexus condenses into a little knot, the supra-oesophageal ganglion. This ganglion cannot do much, if any, thinking; it is rather a steering organ to control the muscles and guide the animal. It is the servant of the locomotive system. Yet it is the beginning of the brain of higher animals, and probably still persists as an infinitesimal portion of our human brain. And all this is the prophecy of a head soon to be developed. An excretory system has appeared to carry off the waste of the muscles and nerves.
In the schematic worm and annelid the reproductive system is simpler, though perhaps equally effective. It takes the excess of nutriment of the body. The muscular system has taken the form of a sack composed of longitudinal and transverse fibres. The perivisceral cavity, formed perhaps by cutting off and enlarging the lateral pouches of the turbellarian digestive system, serves as a very simple but serviceable circulatory system. But in the annelid and all higher forms a special system of tubes has developed to carry the nutriment, and usually oxygen also, needed to keep up the combustion required to furnish the energy in these active organs.
The digestive system has attained its definite form with the appearance of an a.n.a.l opening and the accompanying division of labor and differentiation into fore-, mid-, and hind-intestine.
The digestive and reproductive systems have thus nearly attained their final form. From the higher worms upward the digestive system will improve greatly. Its lining will fold and flex and vastly increase the digestive and absorptive surfaces. The layer of cells which now secrete the digestive fluids will in part be replaced by ma.s.sive glands. Far better means of grasping food than the h.o.r.n.y teeth of annelids will yet appear. But all these changes are inconsiderable compared with the vast advance made by the muscular and nervous systems. Reproduction and digestion are losing their supremacy in the animal body. Their advance and improvement will require but little further attention.
In the annelid especially, and to some extent in the schematic worm, the supra-oesophageal ganglion is relieved in part of the direct control of the muscular fibrils and has become an organ of perception and the seat of government of lower nervous centres. In all higher forms it innervates directly only the princ.i.p.al sense-organs of the head. And at this stage the light-perceiving directive eye has developed into a form-perceiving, eidoscopic organ. The eye was short of range and its images were perhaps rude and imperfect, but it was a visual eye and had vast possibilities.
The animal is taking cognizance of ever more subtle elements in its environment. Perhaps it is not too much to say that the eidoscopic eye first awakened the slumbering animal mind, for its reflex effect upon the supra-oesophageal ganglion cannot be over-estimated. The animal will very soon begin to think.
Between the turbellarian and the annelid many aberrant lines diverged. Some of these attained a comparatively high level and then seemed to meet insuperable obstacles, while others came to an end or turned downward very early. Three of these demanded attention, those leading to mollusks, insects, and vertebrates. And it is interesting to notice that the fundamental difference between these three lines was the skeleton, or perhaps we ought to say it was the habit of life which led to the development of such a skeleton.
The mollusk took to a sluggish, creeping mode of life, under an external purely protective skeleton; the insect to a creeping mode of life, with an external but almost purely locomotive skeleton; the vertebrate kept on swimming and developed an internal locomotive skeleton. And it must already have become clear to you that the destiny of these different lines was fixed not so much directly by the skeleton itself as by its reflex effect in moulding the muscular, and ultimately the nervous, system.
The insects formed their skeleton by thickening the h.o.r.n.y cuticle of the annelid. They transformed the annelid parapodia into legs and developed wings. They attained life in the air. They devoted the muscles of the body largely to the extremities and gained swift locomotion. They have a fair circulatory and an excellent respiratory system. Best of all, they developed a head and a brain by fusing the three anterior ganglia of the body. The insect could and does think. Such a structure ought to lead to great and high results. But actually their possibilities were very limited. They have not progressed markedly during the last geological period.
Their external skeleton was easily attained and brought speedy advantages, which for a time placed them far above all compet.i.tors.
But it limited their size and length of life and opportunities, and finally their intelligence. They remained largely the slaves of instinct. They followed an attractive and exceedingly promising path, but it led to the bottom of a cliff, not to the summit.
The mollusks, clams, and snails took an easier, down-hill road. They formed a sh.e.l.l, and it developed large enough to cover them. It hampered and almost destroyed locomotion and reduced nerve to a minimum. But nerves are nothing but a nuisance anyhow. And why should they move? Food was plenty down in the mud, and if danger threatened, they withdrew into the sh.e.l.l. They stayed down in the mud and let the world go its way. If grievously afflicted by a parasite they produced a pearl--to save themselves from further discomfort. They developed just enough muscle and nervous system to close the sh.e.l.l or drag it a little way; that was all. Digestion and reproduction retained the supremacy. They were fruitful and multiplied, and produced hosts of other clams and snails. The present was enough for them and they had that.
For if the winner in the struggle for existence is the one who gains the most food, the most entire protection against discomfort, danger from enemies or unfavorable surroundings, and the most fruitful and rapid reproduction--and these are all good--then the clam is the highest product of evolution. It never has been surpa.s.sed--I venture to say it never can be--except possibly by the tape-worms. I can never help thinking with what contempt these primitive oysters, if they had had brains enough, would have looked down upon the toiling, struggling, discontented, fighting, aspiring primitive vertebrates.
How they would have wondered why G.o.d allowed such disagreeable, disturbing, unconventional creatures to exist, and thanked him that he had made the world for them, and heaven too, if there be such a place for mollusks. Their road led to the Slough of Contentment.
But even in molluscan history there was a tragic chapter. The squids and cuttle-fishes regained the swimming life, and in their latest forms gave up the protective sh.e.l.l. But its former presence had so modified their structure that any great advance was impossible. It was too late. The sins of the fathers were visited upon the children in the thousandth generation.
The vertebrate developed an internal skeleton. This was necessarily a slow growth, and the type came late to supremacy. The longitudinal muscles are arranged in heavy bands on each side of the back, and the animal swims rapidly. The sense-organs are keen. The brain contains the ganglia of several or many segments and is highly differentiated. It has a special centre of perception, thought, and will; it is an organ of mind. The vertebrate has the physical and mental advantages of large size.
First the definite form and mode of developing a vertebra is attained. Then the vertebral column is perfected. The fins are modified into legs. The lungs increase in size and the heart becomes double. The animal emerges on land; and, with a better supply of oxygen and less loss of heat, all the functions are performed with the highest possible efficiency. First, apparently, amphibia, then reptiles, and finally mammals of enormous size and strength appeared. It looked as if the earth were to be an arena where gigantic beasts fought a never-ending battle of brute force. But these great brutes reproduced slowly, had therefore little power of adaptation, were fitted to special conditions, and when the conditions changed they disappeared. The bird tried once more the experiment of developing the locomotive powers to the highest possible extent. It became a flying machine, and every organ was moulded to suit this life. Every ounce of spare weight was thrown aside, the muscles were wonderfully arranged and of the highest possible efficiency. The body temperature is higher than that of mammals. The whole organization is a physiological high-pressure engine. The sense-organs are perhaps the finest and keenest in the whole animal kingdom. The brain is inferior only to that of mammals.
The experiment could not have been tried under more favorable conditions; it was not a failure, it certainly was not a success when compared with that of mammals.
The possibilities of every system except one had been practically exhausted. Only brain development remained as the last hope of success. Here was an untried line, and the mammals followed it.
During the short tertiary period the brain in many of their genera seems to have increased tenfold. By the arboreal life of the highest forms the hand is developed as the instrument of the thinking brain.
The battle is beginning to become one of wits, and the crown will soon pa.s.s from the strongest to the shrewdest. Mind, not muscle, much less digestion or reproduction, is the goal of the animal kingdom. And we shall see later that the mammalian mode of reproduction and of care of the young led to an almost purely mental and moral advance. For these could have but one logical outcome, family life. And the family is the foundation of society. And family and social life have been the school in which man has been compelled to learn the moral lessons, the application of which has made him what he is.
You must all, I think, have noticed that the different systems of organs succeed one another in a certain definite order; and that each stage from the lowest to the highest is characterized by the predominance of a certain function or group of functions. This sequence of functions is not a deduction but a fact. Place side by side all possible genealogical trees of the animal kingdom, whether founded on comparative anatomy, embryology, palaeontology, or all combined. They will all disclose this sequence of functions arranged in the same order. Let me call your attention to the fact that this order is not due to chance, but rests upon a physiological basis. We might almost claim that if the evolution of man from the single cell be granted, no other order of their occurrence is possible.
The protozoa are mostly, though not purely, nutritive and reproductive. These functions are essential to the existence of the species. Naturally in the early protozoan colonies, and in forms like hydra, these functions predominated. But mere digestive tissue is not enough for digestion. Muscles are needed to draw the food to the mouth, to keep the digestive sack in contact with it, and for other purposes. A little higher they are used to enable the animal to go in search of its food. They are still, however, more or less entirely subservient to digestion. But in the highest worms we are beginning to see signs that muscles are predominating in the body; and we feel that, while mutually helpful, the digestive system exists for the muscles, and these latter are becoming the aim of development. From worms upward there is a marked advance in physical activity and strength. The muscles thicken and are arranged in heavier bands. Skeleton and locomotive appendages and jaws follow in insects and vertebrates. The direct battle of animal against animal, and of strength opposed to strength or activity, becomes ever sharper. The strongest and most active are selected and survive.
And yet this is not the whole truth. Some power of perception is possessed by every animal. But until muscles had developed the nervous system could be of but little practical value. Knowledge of even a great emergency is of little use, if I can do nothing about it. But when the muscles appeared, nerves and ganglion cells were necessary to stimulate and control them. And this highest system holds for a long time a position subordinate to that of the lower muscular organ. Its development seems at first sight extraordinarily slow. Only in insects and vertebrates has it become a centre of instinct and thought. Through the sense-organs it is gaining an ever clearer, deeper, and wider knowledge of its environment. First it is affected only by the lower stimuli of touch, taste, and smell. Then with the development of ear and eye it takes cognizance of ever subtler forces and movements. Memory comes into activity very early.
The animal begins to learn by experience. The brain is becoming not merely a steering but a thinking organ. More and more nervous material is crowded into it and detailed for its work. Wits and shrewdness are beginning to count for something in the battle. Not only the animal with the strongest muscles, but the one with the best brain survives. And thus at last the brain began to develop with a rapidity as remarkable as its long delay. Thus each higher function is called into activity by the next lower, serves this at first, and only later attains its supremacy.
And yet the advance of the different functions is not altogether successive. Muscle and nerve do not wait for digestion and reproduction to show signs of halting before they begin to advance.
They all advance at once. But the progress of reproduction and digestion is most rapid at first, and it appears as if they would outrun the others. But in the ascending series the others follow after, and soon overtake and pa.s.s by them. And these lower functions, when out-marched, do not lag behind, but keep in touch with the others, forming the rear-guard and supply-train of the army. And notice that each organ holds the predominance about as long as it shows the power of rapid improvement. The length of its reign is pretty closely proportional to its capacity of development.
The digestive system reaches that limit early, the muscular system is capable of indefinitely higher complexity, as we see in our hand.
But the muscular system has nearly or quite reached its limit. The body had seen its day of dominance before man arrived on the globe.
But where is the limit to man's mental or moral powers? Every upward step in knowledge, wisdom, and righteousness only opens our eyes to greater heights, before unperceived and still to be attained. These capacities, even to our dim vision, are evidently capable of an indefinite, perhaps infinite, development. What, as yet only partially developed, faculty remains to supersede them? As being capable of an endless development and without a rival, may we not, _must_ we not, consider them as ends in themselves? They are evidently what we are here for. Everything points to a spiritual end in animal evolution. The line of development is from the predominantly material to the predominance of the non-material. Not that the material is to be crowded out. It is to reach its highest development in the service of the mind. The body must be sustained and perfected, but it is not the end. The goal is mind, the body is of subordinate importance.
But if this is true, we must study carefully the development of mind in the animal. The question presses upon us; if there is a sequence of physical functions in animal development, is there not perhaps also a sequence in the development of the mental faculties? What is the crowning faculty of the human mind and how is its fuller development to be attained? Let us pa.s.s therefore to the question of mind in the animal kingdom.
CHAPTER V
THE HISTORY OF MENTAL DEVELOPMENT AND ITS SEQUENCE OF FUNCTIONS
We have sketched hastily the development of the human body. This portion of our history is marked by the successive dominance of higher and higher functions. It is a history treating of successive eras. There is first the period of the dominance of reproduction and digestion, purely vegetative functions, characteristics of the plant just as truly as of the animal. This period extends from the beginning of life up to the time when the annelid was the highest living form yet developed. But in insects and lower vertebrates another system has risen to dominance. This is muscle. The vertebrate no longer devotes all, or the larger part, of its income to digestion and reproduction. If it did, it would degenerate or disappear. The stomach and intestine are improved, but only that they may furnish more abundant nutriment for building and supporting more powerful muscles better arranged. The history of vertebrates is a record of the struggle for supremacy between successive groups of continually greater and better applied muscular power. Here strength and activity seem to be the goal of animal development, and the prize falls to the strongest or most agile. The earth is peopled by huge reptiles, or mammals of enormous strength, and by birds of exceeding swiftness. This portion of our history covers the era of muscular activity.
But these huge brutes are mostly doomed to extinction, and the bird fails of supremacy in the animal kingdom. "The race is not to the swift, nor the battle to the strong." All the time another system has been slowly developing. The complicated nervous system has required ages for its construction and arrangement. Only in the highest mammals does the brain a.s.sert its right to supremacy. But once established on its throne the brain reigns supreme; its right is challenged by no other organ. The possibilities of all the other organs, _as supreme rulers_, have been exhausted. Each one has been thoroughly tested, and its inadequacy proven beyond doubt by actual experiment. These formerly supreme lower organs must serve the higher. The age of man's existence on the globe is, and must remain, the era of mind. For the mind alone has an inexhaustible store of possibilities.
The development of all these systems is simultaneous. From the very beginning all the functions have been represented, all the systems have been gradually advancing. Hydra has a nervous system just as really as man. It has no brain, but it has the potentiality and promise of one, and is taking the necessary steps toward its attainment. But while the development of all is simultaneous, their culmination and supremacy is successive, first stomach and muscle, then brain and mind. That was not first which is spiritual, but that which is natural; and afterward that which is spiritual. But now that the mind has once become supreme, man must live and work chiefly for its higher development. Thus alone is progress possible.
But the word mind calls up before us a long list of powers. And the questions arise, Is one mode and line of mental action just as much the goal of man's development as another? Is man to cultivate the appet.i.te for food and sense gratification just as much as the hunger for righteousness? Or is appet.i.te in the mind like digestion in the body, a function, necessary indeed and once dominant, but no longer fitted for supreme control? Is there in the development of the mental powers or functions just as really a sequence of dominance as in that of the bodily functions? Are there older and lower powers and modes of action, which, though once supreme, must now be rigidly kept down in their proper lower place? Are there lower motives, for which the very laws of evolution forbid us to live, just as truly as they forbid a man's living for stomach or brute strength instead of brain and mind? Are these lower powers merely the foundation on which the higher motives and powers are to rise in their transcendent glory? This is the question which we now must face, and it is of vital importance.
We have come to one of the most important and difficult subjects of zoology. Let us distinctly recognize that it is not our task to explain the origin of mind, or even of a single mental faculty. I shall take for granted what many of you will not admit, that the germs of all man's highest mental powers are present undeveloped in the mind, if you will call it so, of the amoeba. The limits of this course of lectures have required us to choose between alternatives, either to attempt to prove the truth of the theory of evolution, or taking this for granted, to attempt to find its bearings on our moral and religious beliefs. I have chosen the latter course, and here, as elsewhere, will abide by it. I should not have followed such a course if I did not thoroughly believe that man also, in mind as well as body, is the product of evolution. But this is no reason for your accepting these views. You are asked only to judge impartially of the tendencies of the theory. We take for granted, I repeat, that all man's mental faculties are germinally, potentially, present in protoplasm; we seek the history of their development.
We must remember, further, that the science of animal or comparative psychology is yet in its infancy. Even reliable facts are only slowly being sifted and recorded in sufficient numbers to make deductions at all safe. And even of these facts different writers give very different explanations. As Mr. Romanes has well said, "All our knowledge of mental faculties, other than our own, really consists of an inferential interpretation of bodily activities--this interpretation being founded on our subjective knowledge of our own mental activities. By inference we project, as it were, the human pattern of our own mental chromograph on what is to us the otherwise blank screen of another mind." The value and clearness of our inferences will be proportional to the similarity of the animal to ourselves. Thus we can educate many of our higher mammals by a system of rewards and punishments, and we seem therefore to have good reason to believe that fear and joy, anger and desire, certain powers of perception and inference, are in their minds similar to our own. But fear in a fish is certainly a much dimmer apprehension of danger than in us, even if it deserves the name of apprehension.
And the mental state which we call "alarm" in a fly or any lower animal is very difficult to clearly imagine or at all express in terms of our own mind.