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These foot-marks have revealed to us new and unexpected proofs that the air- breathing fauna of the Upper Eocene period in Europe far surpa.s.sed in the number and variety of its species the largest estimate which had previously been formed of it. We may now feel sure that the mammalia, reptiles, and birds which have left portions of their skeletons as memorials of their existence in the solid gypsum const.i.tuted but a part of the then living creation. Similar inferences may be drawn from the study of the whole succession of geological records. In each district the monuments of periods embracing thousands, and probably in some instances hundreds of thousands of years, are totally wanting. Even in the volumes which are extant the greater number of the pages are missing in any given region, and where they are found they contain but few and casual entries of the physical events or living beings of the times to which they relate. It may also be remarked that the subordinate formations met with in two neighbouring countries, such as France and England (the minor Tertiary groups above enumerated), commonly cla.s.sed as equivalents and referred to corresponding periods, may nevertheless have been by no means strictly coincident in date.
Though called contemporaneous, it is probable that they were often separated by intervals of many thousands of years. We may compare them to double stars, which appear single to the naked eye because seen from a vast distance in s.p.a.ce, and which really belong to one and the same stellar system, though occupying places in s.p.a.ce extremely remote if estimated by our ordinary standard of terrestrial measurements.
CALCAIRE SILICIEUX, OR TRAVERTIN INFERIEUR (A.2 AND 3 TABLE 16.1).
This compact siliceous limestone extends over a wide area. It resembles a precipitate from the waters of mineral springs, and is often traversed by small empty sinuous cavities. It is, for the most part, devoid of organic remains, but in some places contains fresh-water and land species, and never any marine fossils. The calcaire siliceux and the calcaire grossier usually occupy distinct parts of the Paris basin, the one attaining its fullest development in those places where the other is of slight thickness. They are described by some writers as alternating with each other towards the centre of the basin, as at Sergy and Osny.
The gypsum, with its a.s.sociated marls before described, is in greatest force towards the centre of the basin, where the calcaire grossier and calcaire silicieux are less fully developed.
GRES DE BEAUCHAMP, OR SABLES MOYENS (A.4 TABLE 16.1).
In some parts of the Paris basin, sands and marls, called the Gres de Beauchamp, or Sables moyens, divide the gypseous beds from the calcaire grossier proper.
These sands, in which a small nummulite (N. variolaria) is very abundant, contain more than 300 species of marine sh.e.l.ls, many of them peculiar, but others common to the next division.
MIDDLE EOCENE FORMATIONS OF FRANCE.
CALCAIRE GROSSIER, UPPER AND MIDDLE (B.1 TABLE 16.1).
The upper division of this group consists in great part of beds of compact, fragile limestone, with some intercalated green marls. The sh.e.l.ls in some parts are a mixture of Cerithium, Cyclostoma, and Corbula; in others Limnea, Cerithium, Paludina, etc. In the latter, the bones of reptiles and mammalia, Palaeotherium and Lophiodon, have been found. The middle division, or calcaire grossier proper, consists of a coa.r.s.e limestone, often pa.s.sing into sand. It contains the greater number of the fossil sh.e.l.ls which characterise the Paris basin. No less than 400 distinct species have been procured from a single spot near Grignon, where they are imbedded in a calcareous sand, chiefly formed of comminuted sh.e.l.ls, in which, nevertheless, individuals in a perfect state of preservation, both of marine, terrestrial, and fresh-water species, are mingled together. Some of the marine sh.e.l.ls may have lived on the spot; but the Cyclostoma and Limnea, being land and fresh-water sh.e.l.ls, must have been brought thither by rivers and currents, and the quant.i.ty of triturated sh.e.l.ls implies considerable movement in the waters.
Nothing is more striking in this a.s.semblage of fossil testacea than the great proportion of species referable to the genus Cerithium (Figures 160 and 161 Chapter 15). There occur no less than 137 species of this genus in the Paris basin, and almost all of them in the calcaire grossier. Most of the living Cerithia inhabit the sea near the mouths of rivers, where the waters are brackish; so that their abundance in the marine strata now under consideration is in harmony with the hypothesis that the Paris basin formed a gulf into which several rivers flowed.
EOCENE FORAMINIFERA.
(FIGURE 219. Calcarina rarispina, Desh.
a. Natural size.
b. Magnified.)
(FIGURE 220. Spirolina stenostoma, Desh.
a. Natural size.
b. Magnified.)
(FIGURE 221. Triloculina inflata, Desh.
a. Natural size.
b. Magnified.)
In some parts of the calcaire grossier round Paris, certain beds occur of a stone used in building, and called by the French geologists "Miliolite limestone." It is almost entirely made up of millions of microscopic sh.e.l.ls, of the size of minute grains of sand, which all belong to the cla.s.s Foraminifera.
Figures of some of these are given in Figures 219 to 221. As this miliolitic stone never occurs in the Faluns, or Upper Miocene strata of Brittany and Touraine, it often furnishes the geologist with a useful criterion for distinguis.h.i.+ng the detached Eocene and Upper Miocene formations scattered over those and other adjoining provinces. The discovery of the remains of Palaeotherium and other mammalia in some of the upper beds of the calcaire grossier shows that these land animals began to exist before the deposition of the overlying gypseous series had commenced.
LOWER CALCAIRE GROSSIER, OR GLAUCONIE GROSSIERE (B.1 TABLE 16.1).
The lower part of the calcaire grossier, which often contains much green earth, is characterised at Auvers, near Pontoise, to the north of Paris, and still more in the environs of Compiegne, by the abundance of nummulites, consisting chiefly of N. laevigata, N. scabra, and N. Lamarcki, which const.i.tute a large proportion of some of the stony strata, though these same foraminifera are wanting in beds of similar age in the immediate environs of Paris.
SOISSONNAIS SANDS, OR LITS COQUILLIERS (B.2 TABLE 16.1).
(FIGURE 222. Nerita conoidea, Lam. Syn. N. Schmidelliana, Chemnitz.)
Below the preceding formation, sh.e.l.ly sands are seen, of considerable thickness, especially at Cuisse-Lamotte, near Compiegne, and other localities in the Soissonnais, about fifty miles N.E. of Paris, from which about 300 species of sh.e.l.ls have been obtained, many of them common to the calcaire grossier and the Bracklesham beds of England, and many peculiar. The Nummulites planulata is very abundant, and the most characteristic sh.e.l.l is the Nerita conoidea, Lam., a fossil which has a very wide geographical range; for, as M. d'Archiac remarks, it accompanies the nummulitic formation from Europe to India, having been found in Cutch, near the mouths of the Indus, a.s.sociated with Nummulites scabra. No less than 33 sh.e.l.ls of this group are said to be identical with sh.e.l.ls of the London clay proper, yet, after visiting Cuisse-Lamotte and other localities of the "Sables inferieurs" of Archiac, I agree with Mr. Prestwich, that the latter are probably newer than the London clay, and perhaps older than the Bracklesham beds of England. The London clay seems to be unrepresented in the Paris basin, unless partially so, by these sands. (d'Archiac Bulletin tome 10 and Prestwich Quarterly Geological Journal 1847 page 377.)
LOWER EOCENE FORMATIONS OF FRANCE.
ARGILE PLASTIQUE (C.2 TABLE 16.1).
At the base of the tertiary system in France are extensive deposits of sands, with occasional beds of clay used for pottery, and called "argile plastique."
Fossil oysters (Ostrea bellovacina) abound in some places, and in others there is a mixture of fluviatile sh.e.l.ls, such as Cyrena cuneiformis (Figure 216), Melania inquinata (Figure 217), and others, frequently met with in beds occupying the same position in the London Basin. Layers of lignite also accompany the inferior clays and sands.
Immediately upon the chalk at the bottom of all the tertiary strata in France there generally is a conglomerate or breccia of rolled and angular chalk-flints, cemented by siliceous sand. These beds appear to be of littoral origin, and imply the previous emergence of the chalk, and its waste by denudation. In the year 1855, the tibia and femur of a large bird equalling at least the ostrich in size were found at Meudon, near Paris, at the base of the Plastic clay. This bird, to which the name of Gastornis Parisiensis has been a.s.signed, appears, from the Memoirs of MM. Hebert, Lartet, and Owen, to belong to an extinct genus.
Professor Owen refers it to the cla.s.s of wading land birds rather than to an aquatic species. (Quarterly Geological Journal volume 12 page 204 1856.)
That a formation so much explored for economical purposes as the Argile plastique around Paris, and the clays and sands of corresponding age near London, should never have afforded any vestige of a feathered biped previously to the year 1855, shows what diligent search and what skill in osteological interpretation are required before the existence of birds of remote ages can be established.
SABLES DE BRACHEUX (C.3 TABLE 16.1).
The marine sands called the Sables de Bracheux (a place near Beauvais), are considered by M. Hebert to be older than the Lignites and Plastic clay, and to coincide in age with the Thanet Sands of England. At La Fere, in the Department of Aisne, in a deposit of this age, a fossil skull has been found of a quadruped called by Blainville Arctocyon primaevus, and supposed by him to be related both to the bear and to the Kinkajou (Cercoleptes). This creature appears to be the oldest known tertiary mammifer.
NUMMULITIC FORMATIONS OF EUROPE, ASIA, ETC.
Of all the rocks of the Eocene period, no formations are of such great geographical importance as the Upper and Middle Eocene, as above defined, a.s.suming that the older tertiary formation, commonly called nummulitic, is correctly ascribed to this group. It appears that of more than fifty species of these foraminifera described by D'Archiac, one or two species only are found in other tertiary formations whether of older or newer date. Nummulites intermedia, a Middle Eocene form, ascends into the Lower Miocene, but it seems doubtful whether any species descends to the level of the London clay, still less to the Argile plastique or Woolwich beds. Separate groups of strata are often characterised by distinct species of nummulite; thus the beds between the lower Miocene and the lower Eocene may be divided into three sections, distinguished by three different species of nummulites, N. variolaria in the upper, N.
laevigata in the middle, and N. planulata in the lower beds. The nummulitic limestone of the Swiss Alps rises to more than 10,000 feet above the level of the sea, and attains here and in other mountain chains a thickness of several thousand feet. It may be said to play a far more conspicuous part than any other tertiary group in the solid framework of the earth's crust, whether in Europe, Asia, or Africa. It occurs in Algeria and Morocco, and has been traced from Egypt, where it was largely quarried of old for the building of the Pyramids, into Asia Minor, and across Persia by Bagdad to the mouths of the Indus. It has been observed not only in Cutch, but in the mountain ranges which separate Scinde from Persia, and which form the pa.s.ses leading to Caboul; and it has been followed still farther eastward into India, as far as eastern Bengal and the frontiers of China.
(FIGURE 223. Nummulites Puschi, D'Archiac. Peyrehorade, Pyrenees.
a. External surface of one of the nummulites, of which longitudinal sections are seen in the limestone.
b. Transverse section of same.)
Dr. T. Thompson found nummulites at an elevation of no less than 16,500 feet above the level of the sea, in Western Thibet. One of the species, which I myself found very abundant on the flanks of the Pyrenees, in a compact crystalline marble (Figure 223) is called by M. D'Archiac Nummulites Puschi. The same is also very common in rocks of the same age in the Carpathians. In many distant countries, in Cutch, for example, some of the same sh.e.l.ls, such as Nerita conoidea (Figure 222), accompany the nummulites, as in France. The opinion of many observers, that the Nummulitic formation belongs partly to the cretaceous era, seems chiefly to have arisen from confounding an allied genus, Orbitoides, with the true Nummulite.
When we have once arrived at the conviction that the nummulitic formation occupies a middle and upper place in the Eocene series, we are struck with the comparatively modern date to which some of the greatest revolutions in the physical geography of Europe, Asia, and Northern Africa must be referred. All the mountain-chains, such as the Alps, Pyrenees, Carpathians, and Himalayas, into the composition of whose central and loftiest parts the nummulitic strata enter bodily, could have had no existence till after the Middle Eocene period.
During that period the sea prevailed where these chains now rise, for nummulites and their accompanying testacea were unquestionably inhabitants of salt water.
Before these events, comprising the conversion of a wide area from a sea to a continent, England had been peopled, as I before pointed out, by various quadrupeds, by herbivorous pachyderms, by insectivorous bats, and by opossums.
Almost all the volcanoes which preserve any remains of their original form, or from the craters of which lava streams can be traced, are more modern than the Eocene fauna now under consideration; and besides these superficial monuments of the action of heat, Plutonic influences have worked vast changes in the texture of rocks within the same period. Some members of the nummulitic and overlying tertiary strata called flysch have actually been converted in the central Alps into crystalline rocks, and transformed into marble, quartz-rock, micha-schist, and gneiss. (Murchison Quarterly Journal of Geological Society volume 5 and Lyell volume 6 1850 Anniversary Address.)
EOCENE STRATA IN THE UNITED STATES.
In North America the Eocene formations occupy a large area bordering the Atlantic, which increases in breadth and importance as it is traced southward from Delaware and Maryland to Georgia and Alabama. They also occur in Louisiana and other States both east and west of the valley of the Mississippi. At Claiborne, in Alabama, no less than 400 species of marine sh.e.l.ls, with many echinoderms and teeth of fish, characterise one member of this system. Among the sh.e.l.ls, the Cardita planicosta, before mentioned (Figure 191), is in abundance; and this fossil and some others identical with European species, or very nearly allied to them, make it highly probable that the Claiborne beds agree in age with the central or Bracklesham group of England, and with the calcaire grossiere of Paris. (See paper by the Author Quarterly Journal of Geological Society volume 4 page 12 and Second Visit to the United States volume 2 page 59.)
Higher in the series is a remarkable calcareous rock, formerly called "the nummulite limestone," from the great number of discoid bodies resembling nummulites which it contains, fossils now referred by A. d'Orbigny to the genus...o...b..toides, which has been demonstrated by Dr. Carpenter to belong to the foraminifera. (Quarterly Journal of Geological Society volume 6 page 32.) That naturalist, moreover, is of opinion that the Orbitoides alluded to (O. Mantelli) is of the same species as one found in Cutch, in the Middle Eocene or nummulitic formation of India.
Above the orbitoidal limestone is a white limestone, sometimes soft and argillaceous, but in parts very compact and calcareous. It contains several peculiar corals, and a large Nautilus allied to N. ziczac; also in its upper bed a gigantic cetacean, called Zeuglodon by Owen. (See Memoir by R.W. Gibbes Journal of Academy of Natural Science Philadelphia volume 1 1847.)
The colossal bones of this cetacean are so plentiful in the interior of Clarke County, Alabama, as to be characteristic of the formation. The vertebral column of one skeleton found by Dr. Buckley at a spot visited by me, extended to the length of nearly seventy feet, and not far off part of another backbone nearly fifty feet long was dug up. I obtained evidence, during a short excursion, of so many localities of this fossil animal within a distance of ten miles, as to lead me to conclude that they must have belonged to at least forty distinct individuals.
(FIGURE 224. Zeuglodon cetoides, Owen. Basilosaurus, Harlan.
Molar tooth, natural size.)
(FIGURE 225. Zeuglodon cetoides, Owen. Basilosaurus, Harlan.
Vertebra, reduced.)
Professor Owen first pointed out that this huge animal was not reptilian, since each tooth was furnished with double roots (Figure 224), implanted in corresponding double sockets; and his opinion of the cetacean nature of the fossil was afterwards confirmed by Dr. Wyman and Dr. R.W. Gibbes. That it was an extinct mammal of the whale tribe has since been placed beyond all doubt by discovery of the entire skull of another fossil species of the same family, having the double occipital condyles only met with in mammals, and the convoluted tympanic bones which are characteristic of cetaceans.